1976-1982 Complutense University, Madrid. MD

1985-1988 Autonomous University, Madrid, and European Molecular Biology Laboratory,
                     Heidelberg. PhD. Studies on the activity of the yeast plasma membrane
                     H+-ATPase. Supervisor: Prof. Ramon Serrano.

1989-1993 German Cancer Research Center. Transgenic models for cervical cancer 
                     caused by HPV16 and 18. Director: Prof. Harald zur Hausen

1994-1997 European Molecular Biology Laboratory. Studies on transcytosis under
                     supervision of Prof. Carlos Dotti

1997-1999 University of Heidelberg. Use of viral vectors to study neuroepithelial
                     differentiation. Director: Prof. Wieland Huttner

2000-date  German Cancer Research Center. Development of therapeutic vaccines for
                     immunotherapy of cervical cancer and other types of cancer.